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Ichthyological Research
An Official Journal of the Ichthyological Society of Japan

Aims and scope:
Ichthyological Research is an official journal of the Ichthyological Society of Japan and is published quarterly in February, May, August, and November. Ichthyological Research primarily publishes research papers on original work, either descriptive or experimental, that advance our understanding of diversity of fishes. Ichthyological Research strives for covering all aspects fish biology, including taxonomy, systematics, evolution, biogeography, ecology, ethology, genetics, morphology, and physiology. Ichthyological Research also publishes timely reviews and commentary on recent progress in active areas of research on fish biology.
Editorial Board
Editor-in-Chief

Hisashi Imamura
Laboratory of Marine Biology and Biodiversity, Faculty of Fisheries Sciences, Hokkaido University
3-1-1 Minato-cho, Hakodate, Hokkaido 041-8611, Japan

Managing Editor

Yoshiaki Kai

Editors

Hiromitsu Endo,
Atsushi Fukui
Masahiro Horinouchi
Tomoyuki Kokita
Kazunori Matsumoto
Hiroyuki Motomura
Edward O. Murdy
Yoshitaka Sakakura
Kunio Sasaki
Hidenobu Yambe
Katsunori Tachihara
Naoki Yagishita
Terry J. Donaldson
Chris Burridge
Whilliam White
Makoto Okamoto
B. Mabel Manjaji-Matsumoto
Yoshinori Taniguchi
Thomas Trunski
Nao Kagawa

Advisory Board

Lee-Shing Fang, Taiwan
Kurt D. Fausch, USA
Jeffrey M. Leis, Australia
Carin Magnhagen, Sweden
Nobuhiko Mizuno, Japan
Jorgen G. Nielsen, Denmark
Mutsumi Nishida, Japan
Lynne R. Parenti, USA
Valentina G. Sideleva, Russia
Edward O. Wiley, USA


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